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Dr Raphael Mrode

Professor of Quantitative Genetics and Genomics
Animal and Veterinary Science

Animal breeding and genetics Team and EGENES

Research interests 

  • Generally interested in data modelling, the application of linear models for data analyses, methods to reduce the dimensions of multi-trait analyses and genetic evaluation of new or novel traits.
  • The incorporation of molecular information in genetic evaluation procedures for traits of economic importance in livestock. This encompasses the use SNPs in computation of genomic breeding values and genomic selection on a within and across breeds basis.
  • Methodologies on the efficient use of sequence data for genetic evaluation.
  • The application of innovative approaches for data capture, analysis and feedback to small holder dairy systems in developing countries
  • Developing and testing innovative genetic and genomic approaches to livestock genetic improvement in small holder systems in developing countries
  • Investigating methods for generating alternative and novel phenotypes in  small dairy systems in developing countries

Selected research projects

  • National genetic and genomic evaluations for daity cattle, beef and sheep                                        Development to support national genetic and genomic evaluations for dairy cattle in the UK
  • Development of national genetic evaluation for resistance to bovine tuberculosis                           Undertaking research and development to implement national genetic and genomic evaluation for TB in the UK
  • Genomic selection in goats                                                                                                                        Developing genetic and genomic evaluations for several traits of economic importance in goats
  • Across Breed Genomic evaluation.                                                                                                                                                               Research into methods that could be employed in genomic evaluations in cross-bred animals
  • The Africa dairy cattle  genetic gain                                                                                                             Research and developments on methods for efficient data capture and feedback systems  for small holder farmers in Africa and  the application of genomic selection


I have been lecturing at the Edinburgh University’s Master’s course on Quantitative genetics and genome analysis since 2005. I have given lectures on mixed linear models and the use of various BLUP models for genetic prediction.  I have invited to give  international courses on quantitative genetics and genomics in Romania, Turkey, Kenya and Ethiopia.

PhD students

  • Alex  Brown                                                                                                                                                          Working on  Across Breed Genomic evaluation.  BBRSC  PhD studentship. Principal supervisor
  • Oluyinka Abejide.                                                                                                                                             Working on Across-country livestock breeding strategies in sub-Saharan Africa. SRUC PhD studentship, Co-supervisor
  • Tomasz Krzyzelewski                                                                                                                                          Working on Novel computing methods for calculating genomic evaluations of livestock. Member of team of Supervisors
  • Ann Maclaren                                                                                                                                                        Worked on “Optimising breeding programmes for the UK sheep industry by the inclusion of  Genotype x Environment (GxE) interaction.’  Co-supervisor. Completed

Selected publications

  • Mucha, S., Mrode, R., I. MacLaren-Lee, M. Coffey, J. Conington (2015). Estimation of genomic breeding values for milk yield in the UK dairy goats. Journal Dairy Science, 98:8201-8208
  • Pickering, N., Chagunda, M., Banos, G., Mrode, R., and McEwan, J., Walls, E., (2015). Genetic parameters for predicted methane production and laser methane detector measurements. Journal of Animal science, 93:11-20
  • Mrode R.A. (2014) Linear models for the prediction of animal breeding values.                                             Third edition, CAB International, Wallingford, Oxon, UK. 360pp
  • McLaren, A., Lambe, N.R., Morgan-Davies, C., Mrode, R.,  Brotherstone, S., Conington, J., Morgan-Davies, J.. and Bunger, L.(2014). Characterisation of terminal sire sheep farm systems, based on a range of environmental factors: a case study in the context of genotype by environment interactions using Charollais lambs. ANIMAL 8:867-876
  • Mrode, R,  T. Pritchard, T., Coffey, M., and Wall, E (2012) Joint estimation of genetic parameters for test-day somatic cell count and mastitis in the United Kingdom. Journal of Dairy Science Journal dairy Science, 95:4618-462

SRUC repository 

Dr Raphael Mrode

Reader in Quantitative Animal Genetics

Address: Animal & Veterinary Sciences, SRUC, Roslin Institute Building, Easter Bush, Midlothian EH25 9RG

Telephone: 0131 651 9312

Fax: 0131 535 3121